Dr. Richard Holt, Bartholomew Lab, Courtesy Research Associate


    Holt Office  Salmon Disease Lab
Phone  541-737-0743
FAX  541-737-0496
Email  holtr@onid.orst.edu
Education Ph.D. Microbiology, OSU

Senior Fish Pathologist, Oregon Department of Fish and Wildlife (Retired).  Forty-two years of experience in the study of fish disease. Previous activities include conducting research in the diagnosis and treatment of fish parasites, pathogens in hatchery fish, detection of pathogens in wild stocks of fish and determining causes of fish losses in Oregon. Principal area of research and publications has been in studies of the yellow pigmented bacterial fish pathogens such as Flavobacterium psychrophilum agent of cold-water disease and F. columnare, agent of columnaris disease. Currently participating on a project to study the occurrence of the myxosporean parasite, Ceratomyxa shasta, and its impact on wild salmonid stocks in the Klamath River watershed. 

SDL Rich Hold


Ray, R. Adam, Holt, R.A. and Bartholomew, J.  2012.  Relationship between temperature and C. shasta-induced mortality in Klamath River salmonids.  J. of Parasitology.  93(3):520-526.

Stocking, R.W., Lorz, H.L., Holt, R.A. and Bartholomew, J.L.  2007.  Surveillance for Ceratomyxa shasta in the Puget Sound Watershed, WA, USA.  J. of Aquatic Animal Hlth. 19:116-120.

Bartholomew, J.L., Lorz, H.V., Atkinson, S.D., Hallett, S.L.  Stevens, D.G., Holt, R.A., Lujan, K. and Amandi, A.  2007.  Evaluation of a management strategy to control the spread of Myxobolus cerebralis in a lower Columbia River tributary.  North American J. of Fisheries Mgmt. 27:542-550.

Stocking, R.W., Holt, R.A., Foott, J.S. and Bartholomew, J.L.  2006.  Spatial and temporal occurrence of the salmonid parasite Ceratomyxa shasta (Myxozoa) in the Oregon-California Klamath River Basin.  J. of Aquatic Animal Health.  18:194-202.

Hallett, S.L, Adkinson, S.D., Holt, R.A., Banner, C.A.  and Bartholomew, J.  2006.  A new Myxozoan from feral goldfish (Carassius auratus).  J. of Parasitology, 92(2):357-363.